There are corner cases where MessageDescriptor.{ClrType,Parser} will return null, and these are now documented. However, normally they *should* be implemented, even for descriptors of for dynamic messages. Ditto FieldDescriptor.Accessor.
We'll still need a fair amount of work to implement dynamic messages, but this change means that the public API will be remain intact.

Additionally, this change starts making use of C# 6 features in the files that it touches. This is far from exhaustive, and later PRs will have more.

Generated code changes coming in the next commit.